This depends on who the "faction creator" is set as in the "descr_strat.txt" file.
The setting defines what the settlement's culture is (shared with the faction creator) and who it will rebel to if it's captured by somebody who is not a member of it's culture group.
Cyrene has Macedon set as its faction creator, so will automatically rebel to the Macedonians if it is captured by a non-Greek faction and then it revolts against its new overlords. This is highly likely to happen in Cyrene, surrounded by Carthaginians and Egpytians.
All cities have a faction creator assigned to them. Often though rebellions to the faction creator are unlikely to occur since they are surrounded by factions of the same culture group as the faction creator. If a faction which is of the same culture group as the creator is holding it, then it defects to rebel control in the case of civil revolts. If the creator has been destroyed the settlement also goes into rebel hands if it revolts.
The faction creator is automatically changed to that of the upgrading faction when the governor's residence is upgraded.
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